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Evolutionary theory: Hamilton's rule OK. Alan Grafen. Naturevolume , pages – () | Download Citation. These are, first, studies that empirically parametrize Hamilton's rule in natural populations and, second, comparative phylogenetic analyses of the genetic. The most extreme examples of altruism in nature come from social insects such Until Hamilton, explanations for the evolution of altruism and.
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Origins of altruism: why Hamilton still rules 50 years on
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